Project proposal details
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The scaling of metabolism with mass in prokaryotes
Project based at
Silwood Park (Imperial)
A major accepted "law" in ecology is that metabolic rate scales with body size (Kleiber's law, Kleiber 1932). In metazoa, metabolic rate increases with body mass with a 3/4 scaling factor, however this power law may be different across other major evolutionary groups (DeLong 2010). Interestingly, recent work (Kremer 2017) seems to show mass scaling relationships in Cyanobacteria departing from previously reported mass scaling across prokaryotes (bacteria and archaea) in general. Are mass scaling laws really universal across major domains of life, or do they differ amongst taxonomic or functional groups (e.g. heterotrophs/autotrophs)? You will test these scaling relationships using our lab's database of microbial growth rates which you will first add to by scanning the literature for microbial cell size information. You will then calculate mass:growth rate scaling relationships across different levels of taxonomic and functional organisation, incorporating temperature normalisation through fitting thermal performance curves for each species. Various models of organism or population rates incorporate mass scaling terms (usually constrained to 0.75), thus understanding the universality of mass scaling relationships is important for better refinement of such modelling work.
1. Kleiber (1932) "Body size and metabolism." Hilgardia
2. DeLong et. al. (2010) "Shifts in metabolic scaling, production, and efficiency across major evolutionary transitions of life." PNAS
3. Kremer et. al. (2017) "Temperature- and size-scaling of phytoplankton population growth rates: Reconciling the Eppley curve and the metabolic theory of ecology." Limnology and Oceanography
Brown et. al. (2004) "Toward a metabolic theory of ecology." Ecology
Uyeda et. a. (2017) "The Evolution of Energetic Scaling across the Vertebrate Tree of Life." The American Naturalist
García et. al. (2016) "The allometry of the smallest: superlinear scaling of microbial metabolic rates in the Atlantic Ocean." The ISME Journal
Ability to program/code in R, Python, or equivalent, and willingness to love our unicellular relatives!
Desk space, computing resources
Project proposal limitations
The project proposer has indicated that there are some limitations to the availability of this project. It may only be available at certain times of year or suit a specific project length. It may also need skills taught to students on a particular course or courses.
Research project proposals are usually part of an active research programme. If supervisors have stated limitations to a proposal, then they are unlikely to have any flexibility. If you are very interested in the topic but have problems with the stated limitations, the supervisor may still be happy to talk to you about other options around the proposal, but you should not expect that any alternative arrangements can be made.
Project length limitations
5 months, 9 months
Available date limitations
Autumn (Sept-Oct), Winter (January), Spring (April-May)
Ecology Evolution and Conservation, Computational Methods in Ecology and Evolution (MSc), Computational Methods in Ecology and Evolution (MRes), Ecological Applications, Ecosystem and Environmental Change MRes